Tom McGrath

Skipper (voice)

Chris Miller

Kowalski (voice)

Christopher Knights

Private (voice)

Conrad Vernon

Rico (voice)

John Malkovich

Dave (voice)

Benedict Cumberbatch

Classified (voice)

Ken Jeong

Short Fuse (voice)

Annet Mahendru

Eva (voice)

Peter Stormare

Corporal (voice)

Andy Richter

Mort (voice)

Danny Jacobs

King Julien (voice)

Werner Herzog

Documentary Filmmaker (voice)

Sean Charmatz

Cricket (voice)

Billy Eichner

New York Reporter (voice)

Stephen Kearin

Pilot / Aquarium Employee (voice)

Ava Acres

Girl at Zoo (voice)

Kelly Cooney Cilella

Mermaid Penguin (voice)

Susan Fitzer

Antarctic Penguin (voice)

Chris Sanders

Antarctic Penguin (voice)

Emily Nordwind

Antarctic Penguin (voice)

Mike Mitchell

Antarctic Penguin (voice)

Hope Levy

Antarctic Penguin / Woman at Aquarium (voice)

Walt Dohrn

Antarctic Penguin (voice)

Jim Pirri

Gondolier (voice)

Elizabeth Pan

Shanghai TV Reporter (voice)

Jeff Fischer

Security System Voice (voice)

Stephen Apostolina

Penguin Prisoner (voice)

Al Rodrigo

Penguin Fan (voice)

Steve Alterman

Aquarium Announcer (voice)

Carter Hastings

Boy at Aquarium (voice)

Cullen McCarthy

Boy at Zoo (voice)

Nicholas Guest

Flight Attendant (voice)

Lynnanne Zager

Computer Voice (voice)