Ray Romano

Manny (voice)

John Leguizamo

Sid (voice)

Denis Leary

Diego (voice)

Seann William Scott

Crash (voice)

Josh Peck

Eddie (voice)

Queen Latifah

Ellie (voice)

Jennifer Lopez

Shira (voice)

Peter Dinklage

Captain Gutt (voice)

Wanda Sykes

Granny (voice)

Keke Palmer

Peaches (voice)

Josh Gad

Louis (voice)

Nick Frost

Flynn (voice)

Aziz Ansari

Squint (voice)

Alain Chabat

Silas (voice)

Kunal Nayyar

Gupta (voice)

Drake

Ethan (voice)

Nicki Minaj

Steffie (voice)

Rebel Wilson

Raz (voice)

Alan Tudyk

Milton / Hunky Siren (voice)

Ester Dean

Female Siren (voice)

Eddie 'Piolin' Sotelo

Uncle Fungus (voice)

Joy Behar

Eunice (voice)

Heather Morris

Katie (voice)

Chris Wedge

Scrat (voice)

Karen Disher

Scratte (voice)

Marieve Herington

Additional Voices (voice)

Allegra Leguizamo

Additional Voices (voice)

Jason Fricchione

Dumb Mammoth / Various (voice)

Ben Gleib

Marshall (voice)

Simon Pegg

Buck (voice)

Alexandra Romano

Meghan (voice)

Patrick Stewart

Ariscratle (voice)

Brian Scott McFadden

Additional Voices (voice)

Edita Brychta

Additional Voices (voice)

Dennis T. Carnegie

Additional Voices (voice)

Aimée Castle

Additional Voices (voice)

Jim Conroy

Additional Voices (voice)

Jason Harris

Additional Voices (voice)

Sondra James

Additional Voices (voice)

Alexa Kahn

Additional Voices (voice)

Bill Lobley

Additional Voices (voice)

Chris Phillips

Additional Voices (voice)

Tara Sands

Additional Voices (voice)

Keith Silverstein

Additional Voices (voice)

Vanessa Lemonides

Additional Voices (voice)

Danny Flaherty

Additional Voices (voice)

Selenis Leyva

Additional Voices (voice)

Ron McClary

Additional Voices (voice)

Peter Pamela Rose

Additional Voices (voice)

Gabriel Rush

Additional Voices (voice)

Randy Thom

Additonal Voices (voice)

Annie Maginnis Tippe

Additional Voices (voice)

Corwin C. Tuggles

Additional Voices (voice)

Tashiana Washington

Additional Voices (voice)

Frank Welker

Giant Crab (voice) (uncredited)